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The Helstrom Ancestor Lizard is a green-skinned lizard; they are named for the city of Helstrom and are known for making great pets. 

The Lilmoth Ancestor Lizard is a black-skinned variant which possesses striking scarlet hues. These lizards are also kept as pets and are known to comfort the pain of others by nuzzling them. They are named for the city of Lilmoth, and some claim that they are the ancestors of the local Bright-Throat Tribe, a coastal tribe of Murkmire Argonians famed for their bright skin coloration. 

Duskfall Ancestor Lizards are black-scaled ancestor lizards valued as gifts from the past by Argonian Sithis worshippers. 

Scholars believe that Argonians were uplifted from these semi-aquatic, tree-dwelling swamp lizards native to the Black Marsh.

The Argonian allegory known as the Parable of Becoming states that the Hist found the humanoid forms of Men and Mer useful, and molded the form of the swamp's lizards after them to create Argonians. 

And in 4E 48, the Argonian Mere-Glim received a vision while on Umbriel of life as a lizard, before the root and the taste of Hist Sap came and transformed both his body and mind. 

It's theorized that ancient Argonian society was more advanced and unified in an age known as the xanmeer period. 

Xanmeers, also known as Stone-Nests, are towering stone citadels and ziggurats built by ancient Argonian civilizations, wherein their priests appeased Sithis in the hopes they would be spared from destruction. Their most infamous gifts to the Void were live blood sacrifices.

How the old ones managed to create such lasting, permanent structures on top of the ever-shifting marshes of their homeland is a secret that has been lost to history. 

Though the ancient Argonians were also adept at metalworking and often used volcanic glass in their weaponry.
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